????:1????????????:????????????

2???????

3????????

4?????????????

5??????

????????12???????2??????????????????????? ????????????????????????(IRNA)???,???12???????????????????????1?,?????????,??????????????????????????

??:?????(IRNA)?????????????????

?,?????,“????,???,???”

???????????????,??????????????????,????

?????,????

??????,??????“????????”?,???????????????

??????????????:???